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Holder Amount Position Size ($ in 1000's)  As of Anson Funds Management LP 4,297,699 $6,318  12/31/2023      SPRO 4,297,699 $6,318  12/31/2023 Vanguard Group Inc. 1,643,757 $2,416  12/31/2023      SPRO 1,643,757 $2,416  12/31/2023 ADAR1 Capital Management LLC 1,067,711 $1,570  12/31/2023      SPRO 1,067,711 $1,570  12/31/2023 Alphabet Inc. 889,979 $1,531  03/31/2024      SPRO 889,979 $1,531  03/31/2024 Atlas Venture Life Science Advisors LLC 1,013,438 $1,490  12/31/2023      SPRO 1,013,438 $1,490  12/31/2023 Murchinson Ltd. 862,700 $1,268  12/31/2023      SPRO 862,700 $1,268  12/31/2023 Renaissance Technologies LLC 547,465 $805  12/31/2023      SPRO 547,465 $805  12/31/2023 BlackRock Inc. 480,599 $706  12/31/2023      SPRO 480,599 $706  12/31/2023 Novo Holdings A S 468,902 $689  12/31/2023      SPRO 468,902 $689  12/31/2023 Geode Capital Management LLC 412,707 $607  12/31/2023      SPRO 412,707 $607  12/31/2023 Ameriprise Financial Inc. 402,963 $592  12/31/2023      SPRO 402,963 $592  12/31/2023 Acadian Asset Management LLC 343,856 $505  12/31/2023      SPRO 343,856 $505  12/31/2023 AWM Investment Company Inc. 333,400 $490  12/31/2023      SPRO 333,400 $490  12/31/2023 Millennium Management LLC 148,857 $219  12/31/2023      SPRO 148,857 $219  12/31/2023 Bridgeway Capital Management Inc. 134,800 $198  12/31/2023      SPRO 134,800 $198  12/31/2023 State Street Corp 130,511 $192  12/31/2023      SPRO 130,511 $192  12/31/2023 DAFNA Capital Management LLC 125,000 $184  12/31/2023      SPRO 125,000 $184  12/31/2023 Squarepoint Ops LLC 122,802 $181  12/31/2023      SPRO 122,802 $181  12/31/2023 Northern Trust Corp 88,881 $131  12/31/2023      SPRO 88,881 $131  12/31/2023 XTX Topco Ltd 65,966 $97  12/31/2023      SPRO 65,966 $97  12/31/2023 DLK Investment Management LLC 31,355 $54  03/31/2024      SPRO 31,355 $54  03/31/2024 GSA Capital Partners LLP 28,319 $49  03/31/2024      SPRO 28,319 $49  03/31/2024 Citadel Advisors LLC 32,274 $47  12/31/2023      SPRO 32,274 $47  12/31/2023 Charles Schwab Investment Management Inc. 28,565 $42  12/31/2023      SPRO 28,565 $42  12/31/2023 Prudential Financial Inc. 28,200 $41  12/31/2023      SPRO 28,200 $41  12/31/2023 Mackenzie Financial Corp 21,462 $32  12/31/2023      SPRO 21,462 $32  12/31/2023 Susquehanna International Group LLP 18,086 $27  12/31/2023      SPRO 18,086 $27  12/31/2023 Morgan Stanley 12,035 $18  12/31/2023      SPRO 12,035 $18  12/31/2023 Dimensional Fund Advisors LP 10,003 $15  12/31/2023      SPRO 10,003 $15  12/31/2023 Texas Yale Capital Corp. 10,000 $15  12/31/2023      SPRO 10,000 $15  12/31/2023 Two Sigma Securities LLC 10,301 $15  12/31/2023      SPRO 10,301 $15  12/31/2023 Russell Investments Group Ltd. 6,400 $9  12/31/2023      SPRO 6,400 $9  12/31/2023 Bank of America Corp DE 5,750 $8  12/31/2023      SPRO 5,750 $8  12/31/2023 Allworth Financial LP 600 $1  03/31/2024      SPRO 600 $1  03/31/2024 Wells Fargo & Company MN 533 $1  12/31/2023      SPRO 533 $1  12/31/2023 Rhumbline Advisers 458 $1  12/31/2023      SPRO 458 $1  12/31/2023 Proequities Inc.   $0  03/31/2024      Put 0 $0  03/31/2024      Call 0 $0  03/31/2024      SPRO 0 $0  03/31/2024 BNP Paribas Arbitrage SA 12 $0  03/31/2024      SPRO 12 $0  03/31/2024 PFS Partners LLC 50 $0  03/31/2024      SPRO 50 $0  03/31/2024 JPMorgan Chase & Co. 236 $0  12/31/2023      SPRO 236 $0  12/31/2023 US Bancorp DE 151 $0  12/31/2023      SPRO 151 $0  12/31/2023 FMR LLC 189 $0  12/31/2023      SPRO 189 $0  12/31/2023 Royal Bank of Canada 315 $0  12/31/2023      SPRO 315 $0  12/31/2023 Tower Research Capital LLC TRC 1 $0  12/31/2023      SPRO 1 $0  12/31/2023 UBS Group AG 329 $0  12/31/2023      SPRO 329 $0  12/31/2023 GWM Advisors LLC 100 $0  12/31/2023      SPRO 100 $0  12/31/2023 Advisor Group Holdings Inc. 97 $0  12/31/2023      SPRO 97 $0  12/31/2023 Qube Research & Technologies Ltd 1 $0  12/31/2023      SPRO 1 $0  12/31/2023 See All: Funds Holding SPRO See Details: Top 10 Hedge Funds Holding SPRO Also See: SPRO Insider Buying Also See: SPRO Holdings Changes

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