There is a long list of 112 institutional holders of INO within the last two holding periods in our database of 13F filers, so we've sorted through to present to you just the top 50 institutional holders of INO by largest to smallest position reported in the institutional investor's latest 13F filing. In the institutional-holders-of-INO table below, the initial column indicates the name of each top institutional holder of INO, with a link to that fund's top holdings. The next columns indicate the amount of INO stock held by each of the institutional holders of INO, followed by the INO position size held by the institution, and finally the date of the quarterly 13F filing period the holdings data represents for that institution.

Holder	Amount	Position Size ($ in 1000's)	As of
Vanguard Group Inc.	19,766,062	$10,081	12/31/2023
INO	19,766,062	$10,081	12/31/2023
D. E. Shaw & Co. Inc.		$7,492	12/31/2023
NOTE	6,500,000	$6,114	12/31/2023
INO	2,701,162	$1,378	12/31/2023
Wolverine Asset Management LLC		$5,329	12/31/2023
NOTE 6	5,690,000	$5,329	12/31/2023
Put	300	$0	12/31/2023
Renaissance Technologies LLC	7,297,379	$3,722	12/31/2023
INO	7,297,379	$3,722	12/31/2023
BlackRock Inc.	3,932,564	$2,006	12/31/2023
INO	3,932,564	$2,006	12/31/2023
GSA Capital Partners LLP	3,796,529	$1,936	12/31/2023
INO	3,796,529	$1,936	12/31/2023
Geode Capital Management LLC	2,871,445	$1,465	12/31/2023
INO	2,871,445	$1,465	12/31/2023
Jacobs Levy Equity Management Inc.	1,149,974	$586	12/31/2023
INO	1,149,974	$586	12/31/2023
Green Alpha Advisors LLC	36,417	$505	03/31/2024
INO	36,417	$505	03/31/2024
Acadian Asset Management LLC	889,002	$452	12/31/2023
INO	889,002	$452	12/31/2023
State Street Corp	871,047	$444	12/31/2023
INO	871,047	$444	12/31/2023
Northern Trust Corp	555,697	$283	12/31/2023
INO	555,697	$283	12/31/2023
Morgan Stanley	541,696	$276	12/31/2023
INO	541,696	$276	12/31/2023
Jane Street Group LLC	461,811	$236	12/31/2023
INO	461,811	$236	12/31/2023
Millennium Management LLC	421,118	$215	12/31/2023
INO	421,118	$215	12/31/2023
Bank of America Corp DE	354,738	$181	12/31/2023
INO	354,738	$181	12/31/2023
Citadel Advisors LLC		$175	12/31/2023
Call	373,800	$191	12/31/2023
Put	32,000	$16	12/31/2023
Invesco Ltd.	333,665	$170	12/31/2023
INO	333,665	$170	12/31/2023
Two Sigma Investments LP	329,088	$168	12/31/2023
INO	329,088	$168	12/31/2023
Cedar Brook Financial Partners LLC	10,876	$151	03/31/2024
INO	10,876	$151	03/31/2024
Susquehanna International Group LLP	289,840	$148	12/31/2023
INO	289,840	$148	12/31/2023
Simplex Trading LLC		$78	12/31/2023
Call	149,400	$76	12/31/2023
INO	11,379	$6	12/31/2023
Put	9,500	$4	12/31/2023
Price T Rowe Associates Inc. MD	150,000	$77	12/31/2023
INO	150,000	$77	12/31/2023
Verition Fund Management LLC	150,045	$77	12/31/2023
INO	150,045	$77	12/31/2023
FMR LLC	146,087	$75	12/31/2023
INO	146,087	$75	12/31/2023
Amundi	139,282	$71	12/31/2023
INO	139,282	$71	12/31/2023
HRT Financial LP	140,633	$71	12/31/2023
INO	140,633	$71	12/31/2023
Tudor Investment Corp Et Al	134,820	$69	12/31/2023
INO	134,820	$69	12/31/2023
Charles Schwab Investment Management Inc.	132,910	$68	12/31/2023
INO	132,910	$68	12/31/2023
Bank of New York Mellon Corp	131,090	$67	12/31/2023
INO	131,090	$67	12/31/2023
Prime Capital Investment Advisors LLC	131,033	$67	12/31/2023
INO	131,033	$67	12/31/2023
Keystone Financial Group	122,737	$63	12/31/2023
INO	122,737	$63	12/31/2023
Two Sigma Advisers LP	117,300	$60	12/31/2023
INO	117,300	$60	12/31/2023
JustInvest LLC	115,361	$59	12/31/2023
INO	115,361	$59	12/31/2023
Pathstone Holdings LLC	110,463	$56	12/31/2023
INO	110,463	$56	12/31/2023
Two Sigma Securities LLC	108,455	$55	12/31/2023
INO	108,455	$55	12/31/2023
Franklin Resources Inc.	98,972	$50	12/31/2023
INO	98,972	$50	12/31/2023
GTS Securities LLC		$42	12/31/2023
Call	71,000	$36	12/31/2023
INO	24,502	$12	12/31/2023
Put	12,200	$6	12/31/2023
Mariner LLC	77,153	$39	12/31/2023
INO	77,153	$39	12/31/2023
Royal Bank of Canada	73,923	$37	12/31/2023
INO	73,923	$37	12/31/2023
Principal Financial Group Inc.	68,349	$35	12/31/2023
INO	68,349	$35	12/31/2023
Dimensional Fund Advisors LP	65,024	$33	12/31/2023
INO	65,024	$33	12/31/2023
XTX Topco Ltd	55,799	$28	12/31/2023
INO	55,799	$28	12/31/2023
UBS Group AG	52,475	$27	12/31/2023
INO	52,475	$27	12/31/2023
Advisor Group Holdings Inc.	52,663	$27	12/31/2023
INO	52,663	$27	12/31/2023
Group One Trading L.P.		$23	12/31/2023
INO	44,360	$23	12/31/2023
Put	4,700	$2	12/31/2023
Call	3,400	$2	12/31/2023
Credit Suisse AG	35,250	$18	12/31/2023
INO	35,250	$18	12/31/2023
Goldman Sachs Group Inc.	35,042	$18	12/31/2023
INO	35,042	$18	12/31/2023
Greenwich Wealth Management LLC	30,580	$16	12/31/2023
INO	30,580	$16	12/31/2023
Mayflower Financial Advisors LLC	30,953	$16	12/31/2023
INO	30,953	$16	12/31/2023